Although amphibians are poor thermoregulators, they do have some control over their body temperature.
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In contrast to mammals and birds, amphibians are unable to produce thermal energy through their metabolic activity, which would allow them to regulate their body temperature independent of the surrounding or ambient temperature. However, the idea that amphibians have no control whatsoever over their body temperature has been proven false because their body temperature does not always correspond to the surrounding temperature. While amphibians are poor thermoregulators, they do exercise control over their body temperature to a limited degree.
Physiological adaptations can assist amphibians in colonizing habitats where extreme conditions prevail. The tolerance range in body temperature represents the range of temperatures within which a species can survive. One species of North American newt is still active when temperatures drop to -2ºC while one South American frog feels comfortable even when temperatures rise to 41ºC-the highest body temperature measured in a free-ranging amphibian. Recently it has been shown that some North American frog and toad species can survive up to five days with a body temperature of -6ºC with approximately one-third of their body fluids frozen. The other tissues are protected because they contain the frost-protective agents glycerin or glucose. Additionally, in many species the tolerance boundaries are flexible and can change as a result of acclimatization (long-term exposure to particular conditions).
Frog species that remain exposed to the sun despite high diurnal (daytime) temperatures exhibit some fascinating modifications in the skin structure that function as morphological adaptations. Most amphibian skin is fully water permeable and is therefore not a barrier against evaporation or solar radiation. The African savanna frog Hyperolius viridiflavus stores guanine crystals in its skin, which enable it to better reflect solar radiation, thus providing protection against overheating. The tree frog Phyllomedusa sauvagei responds to evaporative losses with gland secretions that provide a greasy film over its entire body that helps prevent desiccation (dehydration).
However, behavior is by far the most important factor in thermoregulation. The principal elements in behavioral thermoregulation are basking (heliothermy), heat exchange with substrates such as rock or earth (thigmothermy), and diurnal and annual avoidance behaviors, which include moving to shelter during the day for cooling and hibernating or estivating (reducing activity during cold or hot weather, respectively). Heliothermy is especially common among frogs and toads: it allows them to increase their body temperature by more than 10ºC. The Andean toad Bufo spinulosusexposes itself immediately after sunrise on moist ground and attains its preferred body temperature by this means, long before either ground or air is correspondingly warmed. A positive side effect of this approach is that it accelerates the digestion of the prey consumed overnight, thus also accelerating growth. Thigmothermy is a behavior present in most amphibians, although pressing against the ground serves a dual purpose: heat absorption by conductivity and water absorption through the skin. The effect of thigmothermy is especially evident in the Andean toad during rainfall: its body temperature corresponds to the temperature of the warm earth and not to the much cooler air temperature.
Avoidance behavior occurs whenever physiological and morphological adaptations are insufficient to maintain body temperature within the vital range. Nocturnal activity in amphibians with low tolerance for high ambient temperatures is a typical thermoregulatory behavior of avoidance. Seasonal avoidance behavior is extremely important in many amphibians. Species whose habitat lies in the temperate latitudes are confronted by lethal low temperatures in winter, while species dwelling in arid and semi-arid regions are exposed to long dry, hot periods in summer.
In amphibians hibernation occurs in mud or deep holes away from frost. North of the Pyrenees Mountains, the natterjack toad offers a good example of hibernation, passing the winter dug deep into sandy ground. Conversely, natterjacks in southern Spain remain active during the mild winters common to the region and are instead forced into inactivity during the dry, hot summer season. Summer estivation also occurs by burrowing into the ground or hiding in cool, deep rock crevasses to avoid desiccation and lethal ambient temperatures. Amphibians are therefore hardly at the mercy of ambient temperatures, since by means of the mechanisms described above they are more than able to exercise some control over their body temperature.
题型分类:总结题
文章结构分析:
文章标题预示分类或者机制流程描述结构
首段通过与哺乳动物和鸟类的对比,引入主题:两栖类也能有限度地调节体温。
二段通过具体例子说明,两栖动物具有生理适应调节能力。
三段介绍某些两栖物种的形态适应调节能力,例如皮肤的调节。
四段介绍最重要的温度调节机制:行为。
五段介绍行为调节机制的具体一种,逃避行为的重要性。
六段具体介绍逃避行为的具体方式,冬眠和夏眠的应用。
选项分析:
引导句概括首段结论,预示正确选项应该针对后段调节机制。
Frogs, which选项具体数字二段有出处,但结论(两栖动物能独立于环境温度)与首段结论(exercise control over their body temperature to a limited degree)矛盾,所以错误。
Physical adaptations选项对应二段,正确。
Amphibians can increase选项对应四段,正确。
Sunrise is the time选项针对四段细节,最高级判断没有依据。
Avoidance选项对应五段六段,正确。
Hiberanation选项针对六段细节,概括不准确。
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